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According to this theory one of these extinction pulses occurred at the end of the Guadalupian epoch of the Permian.Marine invertebrates suffered the greatest losses during the P–Tr extinction.During the Triassic, diversity rose rapidly, but disparity remained low.
Because so much biodiversity was lost, the recovery of land-dwelling life took significantly longer than after any other extinction event, However, it is now possible to date the extinction with millennial precision.
Evidence of this was found in samples from south China sections at the P–Tr boundary.
Here, 286 out of 329 marine invertebrate genera disappear within the final two sedimentary zones containing conodonts from the Permian..
Palaeodictyopteroids (insects with piercing and sucking mouthparts) began to decline during the mid-Permian; these extinctions have been linked to a change in flora.
The greatest decline occurred in the Late Permian and was probably not directly caused by weather-related floral transitions.